Corrections and additions to
Epler's 2010 "Water Beetles of Florida"
(last update 28 December 2020)
p. 12.47-12.48 - Hydrophilus
ensifer reported for Florida.
See: Pintar, M.R. & Keller, J.L. 2020. First
record of Hydrophilus ensifer Brullé (Coleoptera:
Hydrophilidae) in the continental United States. The Coleopterists Bulletin
5.116 - Uvarus
sp. 1 has been described as Uvarus
sinofelihelianthus Epler. See Epler, J.H. 2020. A
new species of Uvarus Guignot, 1939 (Coleoptera: Dytiscidae: Hydroporinae:
Bidessini) from Florida, USA. Insecta Mundi 0752: 1-4.
p. 12.40 - Helocombus is now
considered to be part of Cymbiodyta.
See Toussaint, E.F.A. & Short, A.E.Z. 2019. Historical
biogeography of Holarctic Cymbiodyta water scavenger
beetles in the times of Cenozoic land bridge dispersal routes.
Insect Systematics and Diversity 3: 1-7.
p. 16.17 - Prionocyphon
limbatus LeConte has been collected from
FL. Three specimens were collected at Tall Timbers Research
Station in Leon Co. in 1993 (material from FSCA)
p. 12.42 - Hydrobius tumidus is
now Limnohydrobius tumidus
(LeConte). See Short, A.E.Z., Cole, J. & Toussaint,
E.F.A. 2017. Phylogeny, classification and evolution of the
water scavenger beetle tribe Hydrobiusini inferred from morphology
and molecules (Coleoptera: Hydrophilidae). Systematic Entomology
42: 677-691. Of the other two species of Hydrobius
mentioned on p. 12.42 of my FL water beetle manual, melaenus
(Germar) is also now placed in Limnohydrobius; H.
fuscipes (L.) remains in Hydrobius.
p. 5.111 - The species formerly known
as Rhantus calidus
(Fabricius) is now placed in the new genus Meridiorhantus. Thus
it is now known as Meridiorhantus
calidus (Fabricius). See
Balke, M, Hájek, J & Hendrich, L. 2017. Generic
reclassification of species formerly included in Rhantus
Dytiscidae, Colymbetinae). Zootaxa 4258: 91-100.
p. 13.11 - Pronoterus has been
synonymized with Suphisellus.
See Baca, S.M, Toussaint, E.F.A., Miller, K.B. & Short,
A.E.Z. 2017. Molecular phylogeny of the aquatic beetle
family Noteridae (Coleoptera: Adephaga) with an emphasis on data
partitioning strategies. Molecular Phylogenetics and
Evolution 107: 282-292. Thus our single species
of this former genus is now Suphisellus semipunctatus
pp. 6.6.15, 6.17 - Promoresia has been
synonymized with Optioservus.
See Kamite, Y. 2016. Revision
of the genus Optioservus Sanderson, 1953, part 3: The
O. elegans species group. Koleopterologische
Rundschau 86: 205-212. Thus we now have two species of Optioservus
recorded from Florida: O. ovalis (LeConte) and O.
pp. 11.1-11.9 - The species of
Hydrochus have been reviewed for
Mississippi (Worthington, R.J., Hellman, J.L. & Lago,
P.K. 2016. Hydrochidae (Coleoptera) of
Mississippi. Transactions of the American Entomological
Society 142: 167-213. As a result, several of my
letter-designated taxa now have "real" names. H.
sp 1 = H. falsus Hellman; H. sp. 2 = H.
jiawanae Makhan; H. sp. 5 = H. pajnii
Makhan; H. sp. 6 = H. schereri Makhan; H.
sp. 8 = H. jaechi Makhan.
pp. 5.61; 5.110 - Prodaticus has been reduced
to subgeneric status within Hydaticus.
Thus our most common species is now Hydaticus (Prodaticus)
bimarginatus. See Miller &
Bergsten. 2016. Diving Beetles of the
World. Systematics and Biology of the Dytiscidae. Johns
Hopkins University Press. 320 pp.
p. 5.73-5.74 - Coelambus Thomson has been reinstated as a full genus.
Thus two species previously placed in Hygrotus
are now in Coelambus: C. berneri (Young
& Wolfe) and C. nubilus (LeConte).
Note also that "nubilus" was misspelled in the
manual. See Miller, K.B. & Bergsten, J.
2016. Diving Beetles of the World. Systematics and
Biology of the Dytiscidae. Johns Hopkins University
Press. 320 pp.
p. 1.16 - There's a
typo in the first part of couplet 14 leading to
Ptilodactylidae. The last phrase should read "antennal
segments 1 and 2 long, 3 very short". Thanks to Doug Strom
for informing me of my lack of typing skills!
p. 5.78 - The genus Laccomimus
has been established by Toledo & Michat. This genus
replaces Laccodytes Régimbart for the single species known
from Florida, now known as Laccomimus pumilio
(LeConte). See Toledo, M. & Michat, M. 2015. Description
of Laccomimus gen. n. and eleven new species from the
Neotropical region (Coleoptera, Dytiscidae, Laccophilinae).
Zootaxa 3990: 301-354.
pp. 12.8; 12.16, 12.17 - Crenitulus
has been considered a synonym of Anacaena for some
time, but was recently reinstated as a full genus by Fikáček
& Vondráček (2014). Thus Anacaena suturalis (LeConte) becomes Crenitulus
suturalis (LeConte). Anacaena limbata remains the
same, and is still unknown from Florida. For more
information and a corrected key separating the larvae of Anacaena,
Crenitulus and Paracymus download
- Most species of Cyphon from Florida are now placed in Contacyphon.
Zwick, P., Klausnitzer, B. & Ruta, R. 2013. Contacyphon Gozis, 1886
removed from synonymy (Coleoptera: Scirtidae) to accommodate
species so far combined with the invalid name, Cyphon Paykull, 1799.
Entomologische Blätter und Coleoptera 109: 337-353. Note
also that species from the "Cyphon
collaris complex" (none
known to date from Florida) are now placed in the genus Nyholmia. See
Klausnitzer, B. 2013. Monophyletische Artengruppe der
ehemaligen Gattung Cyphon
Paykull, 1799 und Beschreibung von neuen Gattungen (Coleoptera,
Scirtidae). Entomologische Nachrichten und Berichte 57:
12.30 - On p. 12.6 in the key for larval hydrophilids, the third
statement in couplet 12 leading to Enochrus
should read "LEFT mandible
with one inner tooth, RIGHT with two inner teeth".
The figures are oriented correctly. In the diagnosis for
larvae on p. 12.30, the same mistake was repeated. It is the
right mandible that bears two inner teeth; the left mandible bears
one inner tooth. I wonder why it took so long to find this
p. 16.52 - Ora discoidea
Champion has been found in Florida (Collier Co., Immokalee);
it will key to O. texana but
is marked differently.
It can be either mostly tan above or have mostly dark elytra
with a wide pale border, as in the Florida specimen here.
Thanks to Mike Thomas (FSCA) for the photo!
16.22-16.23 - Scirtes goliai Epler
described, replaces Scirtes
sp. 1. See Epler, J.H. 2012. A new species of Scirtes (Coleoptera:
Scirtidae) from southern Florida and the Caribbean. Zootaxa
18.2, 18.7 - parentheses removed from author name for Celina grossula - should be Celina
I had followed an error in Young (1979) by placing "LeConte" in
parentheses. Because LeConte originally described the species grossula in the genus Celina, there are no
parentheses around his name.
pp. 3.1, 3.7 -
To reiterate about the spelling of the name of the weevil Auleutes, it's Auleutes,
not Aleutes or Auletes.
p. 5.38, 5.39,
5.41 - Celina slossonae changed back
to its original spelling - call it Celina slossoni ... again. Note
that even though Mutchler misspelled it as "slossoni" (which is
incorrect; because the species was named for a woman, it should
have been "slossonae"),
Article 33.4 of the ICZN states that such a change is to be termed
an "incorrect subsequent spelling".
p. 5.92 - the
captions for the two beetles are switched; N. p. pullus is on the left,
N. p. floridanus on the
right! Thanks to Dana Denson (RCID) for catching this!
- The genus Stictotarsus is not
known from Florida. Because it is recorded from South
Carolina and there is thus a possibility it may occur here, I
included the genus, represented by the widespread taxon S. griseostriatus (De
Geer), in my key to genera of dytiscid adults (p. 5.26) of
It has long been known that the taxon is notable for its variation
and is most likely a complex of species, especially in Europe. Now
this taxon has been established as a new genus, Boreonectes (Angus, R.B. 2010. Boreonectes gen. n., a new
genus for the Stictotarsus
griseostriatus (De Geer) group of sibling species
(Coleoptera: Dytiscidae), with additional karyosystematic data on
the group. Comparative Cytogenetics 4: 123-131). Thus, the
species’ name is now Boreonectes
griseostriatus (De Geer). Given the variability of
this taxon, it is most likely that at least one of the “variants”
known from North America actually represents an undescribed
species, so stay tuned.
Also note that some other Nearctic Stictotarsus are now included in Boreonectes. These are:
(Clark), B. coelamboides
(Fall), B. dolerosus (Leech),
B. expositus (Fall), B. funereus (Crotch), B. panaminti (Fall)
(tentative placement), B.
spenceri (Leech) and B.
Boreonectes may be
separated from Stictotarsus
by the shining ventral surface of the head behind the eyes,
with only superficial reticulation (closely punctate in Stictotarsus).
p. 1.6 -
2nd column, under 3., line 15 should read “the bulging, white
abdominal venter, ...”
p. 1.8 - Brady Richards (California State University, Chico, CA) was
somehow left off the list of workers who supplied specimens – sorry,
pp. 3.1, 3.7 – Auleutes is
misspelled as “Aleutes”.
p. 5.63 - Couplet 1 has H.
deflatus at 3.8 – 4.7mm but the description has it listed
as 4.2-4.7mm; go with 3.8-4.7 mm.
p. 5.70 - species box: delete space in “inexpectatus”
p. 6.23 – what are termed “basolateral” triangles in couplet 14
should be basomesal triangles
p. 16.14 - under Notes – the “fourth form” referred to in line
1 is not “keyed below”; it should read “discussed below”.